The organelles that are included in the endomembrane system are

The endomembrane system is a group of membrane‑bound organelles that work together to transport and process proteins and lipids within the cell.

Key components of this system include:

1. Endoplasmic reticulum (ER) – a network of membranes where synthesis of proteins (rough ER) and lipids (smooth ER) begins. The ER membrane is continuous with the nuclear envelope, establishing the first link in the system.

2. Golgi complex (Golgi apparatus) – receives vesicles from the ER, modifies their cargo (e.g., glycosylation), and sorts them into new vesicles for delivery to other destinations.

3. Lysosomes – membrane‑bound vesicles that contain hydrolytic enzymes for breaking down macromolecules. They receive material from the Golgi and from endocytosed vesicles, linking them to the same membrane trafficking pathway.

4. Vacuoles – large storage vesicles (especially prominent in plant and fungal cells) that are formed from the same membrane system and function in storage, waste disposal, and maintaining turgor pressure.

These four organelles are all derived from or connected by membrane vesicle traffic, which is why they are classified together as the endomembrane system.

Organelles that are not part of this system, such as mitochondria, ribosomes, and the nucleus, have independent origins and membranes that are not involved in the vesicle‑mediated transport network. Consequently, the correct combination of organelles that belong to the endomembrane system is:

Endoplasmic reticulum, Golgi complex, Lysosomes and Vacuoles.